Recent Publications:

(Senior Researchers: Hershkovitz, May, Sarig, Slon)


  • HalperinSternfeld M, Pokhojaev A, Ghosh M, Rachmiel D, Kannan R, Grinberg I, Asher M, Aviv M, Ma PX, Binderman I,  Sarig R & Adler-Abramovich L. (2023). Immunomodulatory fibrous hyaluronic acid-Fmoc-diphenylalanine-based hydrogel induces bone regeneration. Journal of Clinical Periodontology, 50(2):200-219.
  • Nudel I, Bader-Farraj A, Shpack N, May H & Sarig R (2023). Age estimation of archeological populations using secondary dentin analysis. Journal of Archaeological Science 151:105724.
  • Peer M &  Sarig R. (2023). Color change in teeth due to burning: Spectrophotometric analysis. Forensic Science International 345:111608.
  • Yona S, Medina O, Sarig R & Shvalb N. (2023). An efficient spring model for an integrated orthodontic tooth movement: A verified mathematical model. Applied Sciences 13(8):5013.
  • Fiorenza L, Habashi W, Moggi-Cecchi J, Benazzi S &  Sarig R. (2023). Relationship between interproximal and occlusal wear in Australopithecus africanus and Neanderthal molars. Journal of Human Evolution 183:103423.
  • Elnatan M, Pokhojaev A, Habashi W, Garkun A,  Rittel D & Sarig R. (2023). Investigating the etiology of non-carious cervical lesions: Novel µCT analysis. Journal of Dentistry 136:104615.
  • Francis S, Makoviychuck Y, Chavoinik L, Borgel S, Pokhojaev A, Roul V, Peled N & May H. (2023). A new method for sex estimation based on femoral cross-sectional geometry measurements and its validation using recent and ancient populations. International Journal of Legal Medicine 1-13. ‏
  • Avni HL, Shvalb N, Pokhojaev A, Francis S, Pelleg-Kallevag R, Roul V, Hublin J J H, Rühli F & May H. (2023). Evolutionary roots of the risk of hip fracture in humans. Communications Biology 6.1: 283.
  • Oikonomou A, Asscher Y, May H, Gorin-Rosen Y & Rehren T. (2023). Metal prills in primary glass: A puzzling aspect of the production process. Journal of Archaeological Science-Reports 49: 103977.
  • Kedar E, Koren I, Medlej B & Hershkovitz I. (2023). The Associations between the Maxillary Sinus Volume, Infraorbital Ethmoid Cells, and the Infraorbital Canal: A CT-Based Study. Diagnostics13(23), 3593. 


  • Barzilai, O., Abulafia, T., Shemer, M., May, H., Orbach, M., Frumkin, A., Yeshurun, R., Sarig, R., Porat, N., & Hershkovitz, I. (2022). Rediscovering Geula Cave: A Middle Paleolithic cave site in northern Mt. Carmel, Israel. Quaternary International, 624, 181–197.
  • Bender, O., Megreli, D., Gavish, T., Meyrom, N., Zamir, N., May, H., Sarig, R., & Bar, D. Z. (2022). The hidden secrets of the dental calculus: Calibration of a mass spectrometry protocol for dental calculus protein analysis. International Journal of Molecular Sciences, 23(22), 14387.
  • Ezra, D., Kedar, E., Salame, K., Alperovitch-Najenson, D., & Hershkovitz, I. (2022). Osteophytes on the zygapophyseal (facet) joints of the cervical spine (C3–C7): A skeletal study. Anatomical Record, 305(5), 1065–1072.
  • Garcia, M. H., Benazzi, S., Sarig, R., Zanolli, C., & Fiorenza, L. (2022). Dental macrowear analysis of the Neanderthal mandible Bourgeois-Delaunay 1 (Charente, Southwestern France). Annual Meeting of the European Society for the Study of Human Evolution.
  • Levartovsky, S., Msarwa, S., Reiter, S., Eli, I., Winocur, E., & Sarig, R. (2022). The association between emotional stress, sleep, and awake bruxism among dental students: A sex comparison. Journal of Clinical Medicine, 11(1), 10. 
  • May, H., Sarig, R., Weber, G. W., Fornai, C., Martinon-Torres, M., Bermudez De Castro, J. M., Zaidner, Y., & Hershkovitz, I. (2022). A new Levantine Middle Pleistocene paleodeme: the Nesher Ramla Homo. American Journal of Biological Anthropology, 177, 117.
  • Najafzadeh, A., Garcia, M. H., Benazzi, S., Hublin, J.-J., Kullmer, O., Sarig, R., Sorrentino, R., Vazzana, A., & Fiorenza, L. (2022). Finite Element Analysis of Le Moustier 1 maxillary incisors. Bulletin of the International Association for Paleodontology, 16(2), 209–210.
  • Zohar, I., Alperson-Afil, N., Goren-Inbar, N., Prévost, M., Tütken, T., Sisma-Ventura, G., Hershkovitz, I., & Najorka, J. (2022). Evidence for the cooking of fish 780,000 years ago at Gesher Benot Ya’aqov, Israel. Nature Ecology and Evolution, 6(12), 2016–2028.


  • Zaidner, Y., Centi, L., Prévost, M., Mercier, N., Falguères, C., Guérin, G., ... & Hershkovitz, I. (2021). Middle Pleistocene Homo behavior and culture at 140,000 to 120,000 years ago and interactions with Homo sapiens. Science372(6549), 1429-1433.

  • Hershkovitz, I., May, H., Sarig, R., Pokhojaev, A., Grimaud-Hervé, D., Bruner, E., ... & Zaidner, Y. (2021). A middle pleistocene homo from Nesher Ramla, Israel. Science372(6549), 1424-1428.

  • Hershkovitz, I., Sarig, R., & May, H. (2021). Trends in Ancient Populations’ Osteobiography during the Holocene: the Levantine Perspective. Paléorient, 47(1), 71-82.‏ doi: 10.4000/paleorient.907


  • Floreanova, K., Gilat, E., Koren, I., & May, H. (2020). Ear infection prevalence in prehistoric and historic populations of the southern Levant: A new diagnostic method. International Journal of Osteoarchaeology.
  • Galili, E., Rosen, B., Evron, M. W., Hershkovitz, I., Eshed, V., & Horwitz, L. K. (2020). Israel: Submerged Prehistoric Sites and Settlements on the Mediterranean Coastline—the Current State of the Art. In The Archaeology of Europe’s Drowned Landscapes (pp. 443-481). Springer, Cham.‏  
  • Rothschild, B. M., Tanke, D., Rühli, F., Pokhojaev, A., & May, H. (2020). Suggested case of Langerhans cell Histiocytosis in a cretaceous dinosaur. Scientific Reports10(1), 1-10.‏
  • Peleg, S., Kallevag, R. P., Dar, G., Steinberg, N., Masharawi, Y., & May, H. (2020). New methods for sex estimation using sternum and rib morphology. International Journal of Legal Medicine, 1-12.‏
  • Grimaud-Hervé, D., Albessard-Ball, L., Pokhojaev, A., Balzeau, A., Sarig, R., Latimer, B., ... & Hershkovitz, I. (2020). The endocast of the late Middle Paleolithic Manot 1 specimen, Western Galilee, Israel. Journal of Human Evolution, 102734.‏
  • Abbas, J., Peled, N., Hershkovitz, I., & Hamoud, K. (2020). Pedicle Morphometry Variations in Individuals with Degenerative Lumbar Spinal Stenosis. BioMed Research International.


  • ​Levartovsky, S., Aharonov, O., Emodi Perlman, A., Winocur, E., & Sarig, R. (2019). The effect of tooth wear, age and sex on facial height assessed by soft tissue analysis. Journal of oral rehabilitation.
  • Borgel, S., Latimer, B., McDermott, Y., Sarig, R., Pokhojaev, A., Abulafia, T., ... & May, H. (2019). Early Upper Paleolithic human foot bones from Manot Cave, Israel. Journal of human evolution, 102668.‏
  • Weber, G. W., Hershkovitz, I., Gunz, P., Neubauer, S., Ayalon, A., Latimer, B., ... & May, H. (2019). Before the massive modern human dispersal into Eurasia: A 55,000-year-old partial cranium from Manot Cave, Israel. Quaternary International.‏
  • Sarig, R., Fornai, C., Pokhojaev, A., May, H., Hans, M., Latimer, B., ... & Weber, G. W. (2019). The dental remains from the Early Upper Paleolithic of Manot Cave, Israel. Journal of human evolution, 102648.‏
  • Abbas, J., Peled, N., Hershkovitz, I., & Hamoud, K. (2019). Is Lumbosacral Transitional Vertebra Associated with Degenerative Lumbar Spinal Stenosis?. BioMed Research International2019.‏
  • Feldman, M., Fernández-Domínguez, E., Reynolds, L., Baird, D., Pearson, J., Hershkovitz, I., ... & Bianco, R. A. (2019). Late Pleistocene human genome suggests a local origin for the first farmers of central Anatolia. Nature communications10(1), 1-10.
  • Ezra, D., Slon, V., Kedar, E., Masharawi, Y., Salame, K., Alperovitch‐Najenson, D., & Hershkovitz, I. (2019). The torg ratio of C3–C7 in African Americans and European Americans: A skeletal study. Clinical Anatomy32(1), 84-89.
  • ​Pokhojaev, A., Avni, H., Sella-Tunis, T., Sarig, R., & May, H. (2019). Changes in human mandibular shape during the Terminal Pleistocene-Holocene Levant. Scientific reports9(1), 1-10.
  • Hershkovitz, I. (2019). My hopes for Israel's human-evolution gallery. Nature566(7743), 155-156.
  • Abbas, J., Hamoud, K., Peled, N., & Hershkovitz, I. (2018). Lumbar Schmorl’s nodes and their correlation with spine configuration and degeneration. BioMed research international2018.
  • Hershkovitz, I., Duval, M., Grün, R., Mercier, N., Valladas, H., Ayalon, A., ... & Weinstein-Evron, M. (2018). Response to Comment on “The earliest modern humans outside Africa”. Science362(6413), eaat8964.
  • Ezra, D., Hershkovitz, I., Salame, K., Alperovitch-Najenson, D., Slon, V. (2018). Osteophytes in the cervical vertebral bodies (C3-C7) – Demographical perspectives. Anat Rec. Oct 5.
  • Harney, E., May, H., Shalem, D., Rohland, N., Mallick, S., Lazaridis, I., Sarig, R., Stewardson, K., Nordenfelt, S., Patterson, N., Hershkovitz, I.* & Reich, D*. (2018). Ancient DNA from Chalcolithic Israel reveals the role of population mixture in cultural transformation. Nature communications. Aug 20, 9 (1):1-11.
  • Zohar, I., Dayan, T., Goren, M., Nadel, D., Hershkovitz, I. (2018). Opportunism or aquatic specialization? Evidence of freshwater fish exploitation at Ohalo II – A waterlogged Upper Paleolithic site. PLOS one 13(6) e0198747.
  • Hershkovitz, I., Weber, G. W., Quam, R., Duval, M., Grün, R., Kinsley, L., Ayalon, A., Bar-Matthews, M., Valladas, H., & Weinstein-Evron, M. (2018). The earliest modern humans outside Africa. Science, 359(6374),456-459.
  • Sella-Tunis, T., Pokhojaev, A., Sarig, R., O’Higgins, P., May, H. (2018). Human mandibular shape is associated with masticatory muscle force. Scientific Reports, 8(1), 6042.
  • May, H., Sella-Tunis, T., Pokhojaev, A., Peled, N., & Sarig, R. (2018). Changes in mandible characteristics during the terminal Pleistocene to Holocene Levant and their association with dietary habits. Journal of Archaeological Science: Reports22, 413-419.
  • Pokhojaev, A., Habashi, W., May, H., Schulz-Kornas, E., Shvalb, N., & Sarig, R. (2018). Examination of the Interproximal Wear Mechanism: Facet Morphology and Surface Texture Analysis. Journal of dental research97(13), 1445-1451.
  • Marder, O., Barzilai, O., Abulafia, T., Hershkovitz, I., & Goder-Goldberger, M. (2018). Chrono-cultural considerations of Middle Paleolithic occurrences at Manot Cave (Western Galilee), Israel. In The Middle and Upper Paleolithic Archeology of the Levant and Beyond (pp. 49-63). Springer, Singapore.


  • Sella Tunis, T., Sarig, R., Cohen, H., Medlej, B., Peled, N., May H. (2017). Sex estimation using computed tomography of the mandible International Journal of Legal Medicine. International Journal of Legal Medicine, 1-10,
  • Alex, B. Barzilai, O., Hershkovitz, I., Marder, O., Berna, F., Caracuta, V., Abulafia, T., Davis, L., Goder-Goldberger, M., Lavi, R., Mintz, E., Regev, L., Bar-Yosef Mayer, D., Tejero, J.-M., Yeshurun, R., Ayalon, A., Bar-Matthews, M., Yasur, G., Frumkin, A., Latimer, B., Hans, M. G., & Boaretto, E. (2017). Radiocarbon chronology of Manot Cave, Israel and Upper Paleolithic dispersals: Upper Paleolithic chronology of Manot. Scientific Advances, 3(11), e1701450.​
  • Yeshurun, R., Tejero, J.-M., Barzilai, O., Hershkovitz, I., & Marder, O.  (2017). Upper Palaeolithic bone retouchers from Manot Cave (Israel): A preliminary analysis of a (yet) rare phenomenon in the Levant. The Origins of Bone Tool Technologies, 1-9.
  • Abbas, J., Slon, V., Stein, D., Peled, N., Hershkovitz, I., & Hamoud, K. (2017).In the quest for degenerative lumbar spinal stenosis etiology: the Schmorl's nodes model. BMC Musculoskeletal Disorders18(1), 164.
  • Hershkovitz, I., Latimer, B., Barzilai, O., & Marder, O. (2017). Manot 1 calvaria and recent modern human evolution: an anthropological perspective.” Bulletins et Memoires de la Société d’Anthropologie de Paris, 1-12.
  • Tunis, T. S., Sarig, R., Cohen, H., Medlej, B., Peled, N., & May, H. (2017). Sex estimation using computed tomography of the mandible. International Journal of Legal Medicine, 1-10.
  • Cohen, H., Kugel, C., May, H., Medlej, B., Stein, D., Slon, V., Brosh, T., & Hershkovitz, I. (2017). The effect of impact tool geometry and soft material covering on long bone fracture patterns in children. International Journal of Legal Medicine, 1-11.
  • Cohen, H., Kugel, C., May, H., Medlej, B., Stein, D., Slon, V., & Hershkovitz, I. (2017). The influence of impact direction and axial loading on the bone fracture pattern. Forensic science international277, 197-206.


  • Abbas, J., Slon, V., May, H., Peled, N., Hershkovitz, I., & Hamoud, K. (2016). Paraspinal muscles density: a marker for degenerative lumbar spinal stenosis? BMC Musculoskeletal Disorders17(1), 422.
  • Cohen, H., Kugel, C., May, H., Medlej, B., Stein, D., Slon, V., Hershkovitz, I., & Brosh, T. (2016). The impact velocity and bone fracture pattern: Forensic perspective. Forensic Science International, 266, 54-62.
  • Ezra, D., Mashrawi, Y., Salame, K., Slon, V., Alperovitch-Najenson, D., & Hershkovitz, I. (2016). Demographical aspects in cervical vertebral bodies’ size and shape (c3-c7): a vertebral study. The Spine Journal17(1), 135-142.
  • Stephens, N. B., Kivell, T. L., Gross, T. Pahr, D. H., Lazenby, R. A., Hublin, J. J., Hershkovitz, I., & Skinner, M. (2016). Trabecular architecture in the thumb of Pan and Homo: implications for investigating hand use, loading, and hand preference in the fossil record. American Journal of Physical Anthropology161(4), 603-619.
  • Steinberg, N., Hershkovitz, I., Zeev, N., Rothschild, B., & Siev-Ner, I. (2016). Joint hypermobility and joint range of motion in young dancers. Journal of Clinical Rheumatology, 22(4), 171-178.
  • May, H., & Ruff, C. (2016). Physical burden and lower limb bone structure at the origin of agriculture in the Levant. American Journal of Physical Anthropology161(1), 26-36.
  • Slon, V., Peled, N., Abbas, J., Stein, D., Cohen, H., & Hershkovitz, I. (2016). Vertebral hemangiomas and their correlations with other pathologies. Spine, 41(8), E481-E488.
  • Barzilai, O., Hershkovitz, I., & Marder, O. (2016). The Early Upper Paleolithic Period at Manot Cave, Western Galilee, Israel. Human Evolution, 31(1-2), 85-100.
  • Peleg, S., Dar, G., Steinberg, N., Masharawi, Y., & Hershkovitz, I. (2016). Sacral orientation and Sheuermann’s kyphosis. SpringerPlus, 5(1), 141.
  • Feldman, M., Hershkovitz, I., Sklan, E. H., Bar-Gal, G. K., Pap, I., Szikossy, I., & Rosin-Arbesfeld, R. (2016). Detection of a tumor suppressor gene variant predisposing to colorectal cancer in an 18th century Hungarian mummy. PLoS one11(2), e0147217.
  • Weber, G. W. Fornai, C., Gopher, A., Barkai, R., Sarig, R. & Hershkovitz, I. (2016). The Qesem cave hominin material (part 1): A morphometric analysis of the mandibular premolars and molar. Quaternary International398, 159-174. 
  • Fornai, C., Benazzi, S., Gopher, A., Barkai, R., Sarig, R., Bookstein, F. L., Hershkovitz, I., & Weber, G. W. (2016). The Qesem Cave hominin material (part 2): A morphometric analysis of dm 2-QC2 deciduous lower second molar. Quaternary International398, 175-189.‏
  • Sarig, R., & Tillier, A. M. (2016). Dental wear patterns in early modern humans from Skhul and Qafzeh: A response to Luca Fiorenza and Ottmar Kullmer. HOMO-Journal of Comparative Human Biology67(1), 85-87‏.
  • Tejero, J.-M., Yeshurun, R., Marder, O., Barzilai, O., Hershkovitz, I., Schneller-Pels, N., Bar-Oz, G., Goder-Goldberger, M., & Lavi, R. (2015). The Osseous Industry from Manot Cave (Western Galilee. Israel): Technical and conceptual behaviours of bone and antler exploitation in the Levantine Aurignacian. Quaternary International, 403, 90-106.
  • Sarig, R., Gopher, A., Barkai, R., Rosell, J., Blasco, R., Weber, G. W., Fornai, C., Sella-Tunis, T., & Hershkovitz, I. (2015). How did the Qesem Cave people use their teeth? Analysis of dental wear patterns. Quaternary International398, 136-147.
  • Hershkovitz, I., Weber, G. W., Fornai, C., Gopher, A., Barkai, R., Slon, V., Quam, R., Gabet, Y., & Sarig, R. (2015). New Middle Pleistocene dental remains from Qesem Cave (Israel). Quaternary International398, 148-158.
  • Sarig, R., Hershkovitz, I., Nir, S., May H., & Vardimon, A. D. (2015). Rate and pattern of inter-proximal dental attrition. European Journal of Oral Sciences, 123(4), 276-281.
  • Hardy, K., Radini, A., Buckley, S., Sarig, R., Copeland, L., Gopher, A., & Barkai, R. (2015). Dental calculus reveals potential respiratory irritants and ingestion of essential plant-based nutrients at Lower Palaeolithic Qesem Cave Israel. Quaternary International398, 129-135.
  • Slon, V., Glocke, I., Barkai, R., Gopher, A., Hershkovitz, I., & Meyer, M. (2015). Mammalian mitochondrial capture, a tool for rapid screening of DNA preservation in faunal and undiagnostic remains, and its application to Middle Pleistocene specimens from Qesem Cave (Israel). Quaternary International, 398, 210-218.


  • Hershkovitz, I., Marder, O., Ayalon, A., Bar-Matthews, M., Yasur, G., Boaretto, E., Caracuta, V., Alex, B., Frumkin, A., Goder-Goldberger, M., Gunz, P., Holloway, R. L., Latimer, B., Lavi, R., Matthews, A., Slon, V., Bar-Yosef Mayer, D., Berna, F., Bar-Oz, G., Yeshurun, R., May, H., Hans, M. G., Weber, G. W., & Barzilai. O. (2015). Levantine cranium from Manot Cave (Israel) foreshadows the first European modern humans. Nature, 520(7546), 216-219.
  • Barzilai, O., Marder, O. & Hershkovitz I. (2014). Manot Cave. Seasons 2011-2012. Hadashot Arkheologiot, 126
  • Marder, O., Barzilai, O., Hershkovitz, I., & A. Frumkin. (2013). Karst and prehistory in the western Galilee, emphasizing Manot Cave. In Geological Survey Israel 2013 (pp. 28-36).
  • Marder, O., Alex, B., Ayalon, A., Bar-Matthews, M., Bar-Oz, G., Bar-Yosef Mayer, D., Berna, F., Boaretto, E., Caracuta, V., Frumkin, A., Goder-Goldberger, M., Hershkovitz, I., Latimer, B., Lavi, R., Matthews, A., Weiner, S., Weiss, U., Yas'ur, G., Yeshurun, R., & Barzilai. O. (2013). The Upper Palaeolithic of Manot Cave, Western Galilee, Israel: the 2011–12 excavations. Antiquity, 87(337). 
  • Barzilai, O., Ayalon, A., Bar-Mathews, M., Bar-Oz, G., Boaretto, E., Berna, F., Frumkin, A., Hershkovitz, I., Khalaily, H., Marder, O., Weiner, S., & Yeshurun, R. (2012). Manot Cave: A prehistoric cave site in the western Galilee, Israel. Hadashot Arkheologiot, 124.​


Affiliated with the Faculty of Medicine and the Steinhardt Museum of Natural History

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